group of huge tetrapods that use the tips of their toes or hoofs to walk on

Ungulates (meaning roughly "hoofed" or "hoofed animal") are several groups of mammals most of which use the tips of their toes, usually hoofed, to sustain their whole body weight whilst moving.

Temporal range: Paleocene-Recent,
Possible Late Cretaceous-Present
Donkey in Clovelly, North Devon, England.jpg
Donkey, Equus africanus
Spanish ibex.jpg
Spanish ibex (Capra pyrenaica)
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Clade: Ferungulata
Clade: Ungulata
Linnaeus, 1766
Orders and Clades


  1. Cooper et al. 2014
  2. Welker, F; Collins, MJ; Thomas, JA; Wadsley, M; Brace, S; Cappellini, E; Turvey, ST; Reguero, M; Gelfo, JN; Kramarz, A; Burger, J; Thomas-Oates, J; Ashford, DA; Ashton, PD; Rowsell, K; Porter, DM; Kessler, B; Fischer, R; Baessmann, C; Kaspar, S; Olsen, JV; Kiley, P; Elliott, JA; Kelstrup, CD; Mullin, V; Hofreiter, M; Willerslev, E; Hublin, JJ; Orlando, L; Barnes, I; MacPhee, RD (18 March 2015). "Ancient proteins resolve the evolutionary history of Darwin's South American ungulates" (PDF). Nature. 522 (7554): 81–84. Bibcode:2015Natur.522...81W. doi:10.1038/nature14249. PMID 25799987. Retrieved 27 April 2015.
  3. BURGER, Benjamin J., THE SYSTEMATIC POSITION OF THE SABER-TOOTHED AND HORNED GIANTS OF THE EOCENE: THE UINTATHERES (ORDER DINOCERATA), Utah State University Uintah Basin Campus, Vernal, UT, United States of America, 84078, SVP 2015

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